Histochemical observations in Piper malgassicum (Piperaceae) with a special focus on the epidermis

This is the first contribution about histochemistry of vegetative and reproductive aerial organs in genus Piper L. malgassicum accumulates alkaloids terpenes epidermis underlying layers parenchyma, both leaves, stems anthers. Some idioblasts appear to contain a large amount secondary metabolites. The micro-anatomical analysis showed peculiar secretory structures anthers ovary. Several lipid aggregates, alkaloid droplets calcium oxalate crystals were observed leaves stems, indicating their role defence strategies, mechanical support, pollinators attraction. In anthers, we elaioplasts whose content suggest an alternative indirect function pollination against micro-organisms. Besides, some aggregates surrounded by microtubules, detected recognized as lipotubuloids. tapetum was type. Alkaloids widely distributed plant confirming important biological this type biomolecules its functional range. terpene polyphenol inclusions appeared particularly abundant epidermal layer, whereas close ovule ovary at maturity.